Personal identification

Full name

Susana Carvalho

Citation names

• Carvalho, Susana

Author identifiers

Ciência ID

C91A-A704-6E70

ORCID iD

0000-0003-4542-3720

Languages

Language	Speaking	Reading	Writing	Listening	Peer-review
Portuguese (Mother tongue)
English	Proficiency (C2)	Proficiency (C2)	Proficiency (C2)	Proficiency (C2)	Proficiency (C2)
French	Advanced (C1)	Proficiency (C2)	Intermediate (B1)	Proficiency (C2)	Upper intermediate (B2)
Spanish; Castilian	Proficiency (C2)	Proficiency (C2)	Intermediate (B1)	Proficiency (C2)	Advanced (C1)
Italian	Elementary (A2)	Upper intermediate (B2)	Elementary (A2)	Advanced (C1)	Elementary (A2)

	Degree	Classification
2013/02/13 Concluded	Biological Anthropology (Doutoramento) University of Cambridge Division of Biological Anthropology, United Kingdom
2007/05 Concluded	Mestrado em Biologia e Evolução Humana (Mestrado) Universidade de Coimbra Departamento de Ciências da Terra, Portugal
1997/07 Concluded	História, variante de Arqueologia (Licenciatura) Universidade do Porto Faculdade de Letras, Portugal

Science

	Category Host institution	Employer
2025/02/01 - Current	Invited Principal Investigator (Research)	Centro de Investigação em Biodiversidade e Recursos Genéticos, Portugal
	Associação BIOPOLIS - Rede de Investigação em Biodiversidade e Biologia Evolutiva, Portugal
2024/01 - Current	Coordinating Researcher (Research)	Universidade do Algarve Centro Interdisciplinar de Arqueologia e Evolução do Comportamento Humano, Portugal
	Universidade do Algarve Centro Interdisciplinar de Arqueologia e Evolução do Comportamento Humano, Portugal
2014 - 2015	Postdoc (Research)	The George Washington University Department of Anthropology, United States

Teaching in Higher Education

	Category Host institution	Employer
2021 - 2023	Full Professor (University Teacher)	University of Oxford Institute of Cognitive and Evolutionary Anthropology, United Kingdom
	University of Oxford - Old Road Campus, United Kingdom
2015 - 2021	Associate Professor (University Teacher)	University of Oxford Institute of Cognitive and Evolutionary Anthropology, United Kingdom
	University of Oxford - Old Road Campus, United Kingdom

Positions / Appointments

	Category Host institution	Employer
2024/01 - Current	Organic Unit Sub-director	Gorongosa National Park, Mozambique
	Gorongosa National Park, Mozambique

Grant

	Designation	Funders
2024/07 - 2028	A comparative framework for hominin phylogeography: genetic structure of baboon species (Papio sp.) in the southernmost Great Rift Valley 1240657 Co-Principal Investigator (Co-PI) Pontificia Universidad Católica de Chile, Chile	Fondo Nacional de Desarrollo Científico Tecnológico y de Innovación Tecnológica Ongoing
2021/09/01 - 2023/08/31	Deep(er) Learning of our Behavioural Evolution: Tracking Bipedalism and Predation in Terrestrial Primates MF21\210041 Principal investigator University of Oxford School of Anthropology and Museum Ethnography, United Kingdom	The British Academy
2020 - 2021	Ecology and the deep roots of our African origins AE02.01 Principal investigator University of Oxford - Old Road Campus, United Kingdom National Museums of Kenya, Kenya	University of Oxford - Old Road Campus Concluded
2020 - 2021	Ecology and the deep roots of our African origins AE02.01	University of Oxford - Old Road Campus
2019 - 2021	Monkeying around, complexity, cognition and culture in primate play NIFBA19\191052 Co-Principal Investigator (Co-PI) University of Oxford - Old Road Campus, United Kingdom	The British Academy Concluded
2019 - 2020	Searching for our deep African roots south of the equator NGS-57285R-19 Principal investigator University of Oxford - Old Road Campus, United Kingdom Gorongosa National Park, Mozambique	National Geographic Society Concluded
2018 - 2019	Primate Adaptations to Complex environments and implications for human evolution Gr. CONF-789 Co-Principal Investigator (Co-PI) University of Oxford - Old Road Campus, United Kingdom Gorongosa National Park, Mozambique	Wenner-Gren Foundation for Anthropological Research Concluded
2016 - 2019	Technological Origins: Environmental and Behavioral Context of the Earliest Tool Users Award VF1\103623 Co-Principal Investigator (Co-PI) The George Washington University, United States University of Oxford - Old Road Campus, United Kingdom	National Science Foundation Concluded
2008/05/01 - 2012/04/30	CHIMPANZEE ARCHAEOLOGY: BRIDGING NATURAL AND SOCIAL SCIENCES IN SEEKING THE EVOLUTIONARY ORIGINS OF TECHNOLOGY SFRH/BD/36169/2007 University of Cambridge, United Kingdom	Fundação para a Ciência e a Tecnologia Concluded

Contract

	Designation	Funders
2019/01/01 - 2019/12/31	Interdisciplinary Center for Archaeology and Evolution of Human Behaviour UID/ARQ/04211/2019 Universidade do Algarve, Portugal Universidade do Algarve Centro Interdisciplinar de Arqueologia e Evolução do Comportamento Humano, Portugal	Fundação para a Ciência e a Tecnologia Concluded
2019/01/01 - 2019/12/31	Centre for Functional Ecology UID/BIA/04004/2019 Universidade de Coimbra Centre for Functional Ecology - Science for People & the Planet, Portugal Universidade de Coimbra, Portugal	Fundação para a Ciência e a Tecnologia Concluded
2011/01/01 - 2012/12/31	Strategic Project - UI 283 - 2011-2012 PEst-OE/SADG/UI0283/2011 Universidade de Coimbra, Portugal Universidade de Coimbra Centro de Investigação em Antropologia e Saúde, Portugal	Fundação para a Ciência e a Tecnologia Concluded

Publications

Book chapter	Carvalho, Susana. "Susana Carvalho". In Humans. United States: Columbia University Press, 2023. Pereira, Telmo; Paixão, Eduardo; Carvalho, Vânia Sofia; Carvalho, Susana; Gomes, Rui Telmo. "Raw material diversity, distribution and acquisition in the river Lis basin, central Portugal". In Raw materials exploitation in Prehistory: sourcing, processing and distribution, 16-29. Cambridge: Cambridge Scholars Publishing, 2017. Rolian Campbell; Carvalho, Susana. "Tool use and manufacture in the last common ancestor of Pan and Homo". In Chimpanzees and Human Evolution., 602-644. Harvard, United States: Harvard University Press, 2017. Published Carvalho, Susana. "Extensive surveys of chimpanzee stone tools: From the telescope to the magnifying glass.". In The Chimpanzees of Bossou and Nimba: A Cultural Primatology., 145-156. Tokyo, Japan: Springer, 2011. Published Carvalho, Susana. "Diecké Forest, Guinea: Delving into chimpanzee behavior using stone tool surveys". In The Chimpanzees of Bossou and Nimba: A Cultural Primatology., 301-312. Tokyo, Japan: Springer, 2011. Published Dora Biro; Carvalho, Susana; Matsuzawa, T.. Corresponding author: Dora Biro. "Tools, traditions and technologies: Interdisciplinary approaches to chimpanzee nut-cracking.". In The Mind of the Chimpanzee: Ecological and Experimental Perspectives, 141-155. Chicago, United States: University of Chicago Press, 2010. Published
Conference abstract	Lopes, Célia; Marques, Richard; Campanacho, Vanessa; Carvalho, Susana; Carvalho, Vânia; Cruz, Cristina Barroso; Curate, Francisco; et al. "GEEvH e a Comunicação de Ciência: existe espaço para o ensino da Evolução Humana?". Paper presented in SciComPt 2018, Figueira de Castelo Rodrigo, 2018. Marques, Richard; Campanacho, Vanessa; Carvalho, Susana; Carvalho, Vânia; Cruz, Cristina Barroso; Curate, Francisco; Gonçalves, David; et al. "GEEvH, associativismo científico e comunicação de ciência: 10 anos a divulgar Evolução Humana". Paper presented in 2º Encontro de História da Ciência no Ensino, Coimbra, 2017. Published
Conference paper	REGALA, FREDERICO; Pinto, Maria José da Rosa; Paulo, Luís Meira; Bobe, Rene; Aldeias, Vera; Archer, Will; Braun, David R.; et al. Corresponding author: REGALA, FREDERICO. "The caves of Gorongosa National Park, Mozambique". Paper presented in 18th International Congress of Speleology, Savoie Mont Blanc, 2022. Published
Conference poster	Cruz, Cristina Barroso Silva da; Cunha, Eugénia; Assis, Sandra; Marques, Rui; Carvalho, Vânia; Sousa, Ana Cristina; Nogueiro, Inês; et al. "A interdisciplinaridade em Antropologia: o caso da Evolução Humana". Paper presented in Congresso da Associação Portuguesa de Antropologia, 2006.
Encyclopedia entry	Carvalho, Susana; Beardmore-Herd, Megan. Corresponding author: Carvalho, Susana. "Technological Origins: Primate Perspectives and Early Hominin Tool Use in Africa". In Oxford Research Encyclopedia of African History. Oxford University Press, 2019. http://dx.doi.org/10.1093/acrefore/9780190277734.013.75. 10.1093/acrefore/9780190277734.013.75 Carvalho, Susana; Almeida-Warren, Katarina. "Primate Archaeology". In Encyclopedia of Animal Behavior. Elsevier, 2019. Published
Journal article	Megan Beardmore-Herd; Meredith S. Palmer; Kaitlyn M. Gaynor; Susana Carvalho. "Effects of an Extreme Weather Event on Primate Populations". American Journal of Biological Anthropology (2025): https://doi.org/10.1002/ajpa.25049. 10.1002/ajpa.25049 Braun, D. R.; Carvalho, Susana; Kaplan, R. S.; Beardmore-Herd, Megan; Plummer, T.; Biro, D.; Matsuzawa, T.. "Stone selection by wild chimpanzees shares patterns with Oldowan hominins". (2025): http://hdl.handle.net/10400.1/26757. 10.1016/j.jhevol.2024.103625 Wang, Xueye; Bocksberger, Gaëlle; Arandjelovic, Mimi; Agbor, Anthony; Angedakin, Samuel; Aubert, Floris; Ayimisin, Emmanuel Ayuk; et al. "Strontium isoscape of sub-Saharan Africa allows tracing origins of victims of the transatlantic slave trade". Nature Communications 15 1 (2024): https://doi.org/10.1038/s41467-024-55256-0. 10.1038/s41467-024-55256-0 Berdugo, S.; Cohen, E.; Davis, A. J.; Matsuzawa, T.; Carvalho, S.. "Reliable long-term individual variation in wild chimpanzee technological efficiency". Nature Human Behaviour (2024): https://doi.org/10.1038/s41562-024-02071-8. Published • 10.1038/s41562-024-02071-8 Luncz, Lydia V.; Slania, Nora E.; Almeida-Warren, Katarina; Carvalho, Susana; Falótico, Tiago; Malaivijitnond, Suchinda; Arroyo, Adrián; de la Torre, Ignacio; Proffitt, Tomos. "Tool skill impacts the archaeological evidence across technological primates". Scientific Reports 14 1 (2024): https://doi.org/10.1038/s41598-024-67048-z. Published • 10.1038/s41598-024-67048-z Eppley, Timothy M.; Reuter, Kim E.; Sefczek, Timothy M.; Tinsman, Jen; Santini, Luca; Hoeks, Selwyn; Andriantsaralaza, Seheno; et al. "Tropical field stations yield high conservation return on investment". Conservation Letters 17 2 (2024): https://doi.org/10.1111/conl.13007. Published • 10.1111/conl.13007 Elodie Freymann; João d'Oliveira Coelho; Catherine Hobaiter; Michael A. Huffman; Geresomu Muhumuza; Klaus Zuberbühler; Susana Carvalho. "Applying collocation and APRIORI analyses to chimpanzee diets: Methods for investigating nonrandom food combinations in primate self-medication". American Journal of Primatology (2024): https://doi.org/10.1002/ajp.23603. 10.1002/ajp.23603 Baehren, Lucy; Carvalho, Susana. "Conceptual Archaeology of a Temporal Place: Albert and Kessler (1976) Applied to Chacma Baboons (Papio ursinus)". Journal of Social Sciences 20 1 (2024): 1-11. https://doi.org/10.3844/jssp.2024.1.11. Published • 10.3844/jssp.2024.1.11 Negash, Enquye W.; Alemseged, Zeresenay; Barr, W. Andrew; Behrensmeyer, Anna K.; Blumenthal, Scott A.; Bobe, René; Carvalho, Susana; et al. "Modern African ecosystems as landscape-scale analogues for reconstructing woody cover and early hominin environments". (2024): http://hdl.handle.net/10400.1/26503. 10.1016/j.jhevol.2024.103604 Freymann, E.; Badihi, G.; Hobaiter, C.; Huffman, M. A.; Muhumuza, G.; Orbell, S.; Sempebwa, D.; et al. "The adaptive role of bark in the diet of budongo chimpanzees (Pan troglodytes schweinfurthii)". (2024): http://hdl.handle.net/10400.1/25993. 10.1007/s10764-024-00445-3 Freymann, Elodie; Carvalho, Susana; Garbe, Leif A.; Ghazhelia, Dinda Dwi; Hobaiter, Catherine; Huffman, Michael A.; Muhumuza, Geresomu; et al. "Pharmacological and behavioral investigation of putative self-medicative plants in budongo chimpanzee diets". (2024): http://hdl.handle.net/10400.1/26020. 10.1371/journal.pone.0305219 Michelle Bezanson; Liliana Cortés-Ortiz; Júlio César Bicca-Marques; Ramesh Boonratana; Susana Carvalho; Marina Cords; Stella de la Torre; et al. "News and Perspectives: Words matter in primatology". Primates (2023): https://doi.org/10.1007/s10329-023-01104-6. 10.1007/s10329-023-01104-6 Alejandra Pascual-Garrido; Susana Carvalho; Katarina Almeida-Warren. "Primate archaeology 3.0". American Journal of Biological Anthropology (2023): https://doi.org/10.1002/ajpa.24835. 10.1002/ajpa.24835 Bobe, René; Aldeias, Vera; Alemseged, Zeresenay; Anemone, Robert L.; Archer, Will; Aumaître, Georges; Bamford, Marion K.; et al. "The first Miocene fossils from coastal woodlands in the southern East African Rift". iScience 26 9 (2023): 107644. http://dx.doi.org/10.1016/j.isci.2023.107644. 10.1016/j.isci.2023.107644 Daniel P. Schofield; Gregory F. Albery; Josh A. Firth; Alexander Mielke; Misato Hayashi; Tetsuro Matsuzawa; Dora Biro; Susana Carvalho. "Automated face recognition using deep neural networks produces robust primate social networks and sociality measures". Methods in Ecology and Evolution (2023): https://doi.org/10.1111/2041-210X.14181. 10.1111/2041-210X.14181 Villaseñor, Amelia; Uno, Kevin T.; Kinyanjui, Rahab N.; Behrensmeyer, Anna K.; Bobe, René; Advokaat, Eldert L.; Bamford, Marion; et al. "Pliocene hominins from East Turkana were associated with mesic environments in a semiarid basin". Journal of Human Evolution 180 (2023): 103385. http://dx.doi.org/10.1016/j.jhevol.2023.103385. 10.1016/j.jhevol.2023.103385 Koops, Kathelijne; Biro, Dora; Matsuzawa, Tetsuro; McGrew, William C.; Carvalho, Susana. "Appropriate knowledge of wild chimpanzee behavior (‘know-what’) and field experimental protocols (‘know-how’) are essential prerequisites for testing the origins and spread of technological behavior. Response to “Unmotivated subjects cannot provide interpretable data and tasks with sensitive learning periods require appropriately aged subjects” by C. Tennie and J. Call". Animal Behavior and Cognition 10 2 (2023): 163-168. http://dx.doi.org/10.26451/abc.10.02.05.2023. 10.26451/abc.10.02.05.2023 Lüdecke, Tina; Leichliter, Jennifer N.; Aldeias, Vera; Bamford, Marion K.; Biro, Dora; Braun, David R.; Capelli, Cristian; et al. "Carbon, nitrogen, and oxygen stable isotopes in modern tooth enamel: A case study from Gorongosa National Park, central Mozambique". Frontiers in Ecology and Evolution 10 (2022): http://dx.doi.org/10.3389/fevo.2022.958032. 10.3389/fevo.2022.958032 Luncz, Lydia V.; Braun, David R.; Marreiros, Joao; Bamford, Marion; Zeng, Chen; Pacome, Serge Soiret; Junghenn, Patrick; et al. "Chimpanzee wooden tool analysis advances the identification of percussive technology". iScience 25 11 (2022): 105315. http://dx.doi.org/10.1016/j.isci.2022.105315. Published • 10.1016/j.isci.2022.105315 Lucy Baehren; Susana Carvalho. "Yet Another Non-Unique Human Behaviour: Leave-Taking in Wild Chacma Baboons (Papio ursinus)". Animals (2022): https://doi.org/10.3390/ani12192577. 10.3390/ani12192577 Hammond, Philippa; Lewis-Bevan, Lynn; Biro, Dora; Carvalho, Susana. "Risk perception and terrestriality in primates: A quasi-experiment through habituation of chacma baboons (Papio ursinus) in Gorongosa National Park, Mozambique". American Journal of Biological Anthropology 179 1 (2022): 48-59. http://dx.doi.org/10.1002/ajpa.24567. Published • 10.1002/ajpa.24567 Santander, Cindy; Molinaro, Ludovica; Mutti, Giacomo; Martínez, Felipe I.; Mathe, Jacinto; Ferreira da Silva, Maria Joana; Caldon, Matteo; et al. "Genomic variation in baboons from central Mozambique unveils complex evolutionary relationships with other Papio species". BMC Ecology and Evolution 22 1 (2022): http://dx.doi.org/10.1186/s12862-022-01999-7. 10.1186/s12862-022-01999-7 Koops, Kathelijne; Arandjelovic, Mimi; Hobaiter, Catherine; Kalan, Ammie; Luncz, Lydia; Musgrave, Stephanie; Samuni, Liran; Sanz, Crickette; Carvalho, Susana. "Chimpanzee culture in context". Physics of Life Reviews 44 (2022): 77-80. http://dx.doi.org/10.1016/j.plrev.2022.12.003. Published • 10.1016/j.plrev.2022.12.003 Carvalho, Susana; Wessling, Erin G.; Abwe, Ekwoge E.; Almeida-Warren, Katarina; Arandjelovic, Mimi; Boesch, Christophe; Danquah, Emmanuel; et al. "Using nonhuman culture in conservation requires careful and concerted action". Conservation Letters 15 2 (2022): http://dx.doi.org/10.1111/conl.12860. Published • 10.1111/conl.12860 João d’Oliveira Coelho; Robert L. Anemone; Susana Carvalho. "Unsupervised learning of satellite images enhances discovery of late Miocene fossil sites in the Urema Rift, Gorongosa, Mozambique". PeerJ (2021): https://doi.org/10.7717/peerj.11573. 10.7717/peerj.11573 Almeida-Warren, K.; Braun, D.R.; Carvalho, S.; Almeida-Warren, Katarina; Braun, David R.; Carvalho, Susana. "The DistoX2: A methodological solution to archaeological mapping in poorly accessible environments". Journal of Archaeological Science: Reports 35 (2021): http://www.scopus.com/inward/record.url?eid=2-s2.0-85098199022&partnerID=MN8TOARS. 10.1016/j.jasrep.2020.102688 René Bobe; Felipe I. Martínez; Susana Carvalho. "Primate adaptations and evolution in the Southern African Rift Valley". Evolutionary Anthropology: Issues, News, and Reviews (2020): https://doi.org/10.1002/evan.21826. 10.1002/evan.21826 Bobe, René; Manthi, Fredrick Kyalo; Ward, Carol V.; Plavcan, J. Michael; Carvalho, Susana. "The ecology of Australopithecus anamensis in the early Pliocene of Kanapoi, Kenya". Journal of Human Evolution 140 (2020): 102717. http://dx.doi.org/10.1016/j.jhevol.2019.102717. 10.1016/j.jhevol.2019.102717 da Silva, Maria Ferreira; Tralma, Paula; Martinez, Felipe I.; Capelli, Cristian; Carvalho, Susana. "Investigating Papio sp. Genetic Variation and Gene Flow in Gorongosa, Mozambique, to Improve the Understanding of Introgression or Admixture Processes Between Baboons Species in the Top End of the Great Rift Valley". Folia Primatologica (2020): https://publons.com/wos-op/publon/35167758/. Daniel Schofield; Arsha Nagrani; Andrew Zisserman; Misato Hayashi; Tetsuro Matsuzawa; Dora Biro; Susana Carvalho. "Chimpanzee face recognition from videos in the wild using deep learning". Science Advances 5 9 (2019): https://doi.org/10.1126/sciadv.aaw0736. 10.1126/sciadv.aaw0736 Martinez, Felipe I.; Capelli, Cristian; Ferreira da Silva, Maria J.; Aldeias, Vera; Alemseged, Zeresenay; Archer, William; Bamford, Marion; et al. "A missing piece of the Papio puzzle: Gorongosa baboon phenostructure and intrageneric relationships". Journal of Human Evolution 130 (2019): 1-20. http://dx.doi.org/10.1016/j.jhevol.2019.01.007. 10.1016/j.jhevol.2019.01.007 Gonçalves, André; Carvalho, Susana. "Death among primates: a critical review of non-human primate interactions towards their dead and dying". Biological Reviews 94 4 (2019): 1502-1529. http://dx.doi.org/10.1111/brv.12512. 10.1111/brv.12512 Thompson, Jessica C.; Carvalho, Susana; Marean, Curtis W.; Alemseged, Zeresenay. "Origins of the Human Predatory Pattern: The Transition to Large-Animal Exploitation by Early Hominins". Current Anthropology 60 1 (2019): 1-23. http://dx.doi.org/10.1086/701477. 10.1086/701477 Habermann, Jörg M.; Alberti, Matthias; Aldeias, Vera; Alemseged, Zeresenay; Archer, Will; Bamford, Marion; Biro, Dora; et al. Corresponding author: Habermann, Jörg M.. "Gorongosa by the sea: First Miocene fossil sites from the Urema Rift, central Mozambique, and their coastal paleoenvironmental and paleoecological contexts". Palaeogeography, Palaeoclimatology, Palaeoecology 514 (2019): 723-738. http://dx.doi.org/10.1016/j.palaeo.2018.09.032. 10.1016/j.palaeo.2018.09.032 Bobe, René; Carvalho, Susana. "Hominin diversity and high environmental variability in the Okote Member, Koobi Fora Formation, Kenya". Journal of Human Evolution 126 (2019): 91-105. http://dx.doi.org/10.1016/j.jhevol.2018.10.012. 10.1016/j.jhevol.2018.10.012 Bobe, René; Carvalho, Susana. "The decline of Africa's largest mammals". Science 362 6417 (2018): 892-893. http://dx.doi.org/10.1126/science.aav6883. 10.1126/science.aav6883 Lucy Tibble; Susana Carvalho. "Rethinking the evolution of property and possession: A review and methodological proposition". Evolutionary Anthropology: Issues, News, and Reviews (2018): https://doi.org/10.1002/evan.21748. 10.1002/evan.21748 Mendonça, Renata S.; Dahl, Christoph D.; Carvalho, Susana; Matsuzawa, Tetsuro; Adachi, Ikuma. "Touch-screen-guided task reveals a prosocial choice tendency by chimpanzees (Pan troglodytes)". PeerJ 6 (2018): e5315. https://doi.org/10.7717/peerj.5315. 10.7717/peerj.5315 Carvalho, Susana; Aldeias, Vera; Alemseged, Zeresenay; Archer, William; Bamford, Marion; Biro, Dora; Bobe, René; et al. Corresponding author: Carvalho, Susana. "Gorongosa National Park and the biogeography of human origins in the Mio-Pliocene". Proceedings of the European Society for the Study of Human Evolution 36 (2017): Published Carvalho, S.. "Standing on the shoulders of giants: The contribution of Cláudia Sousa for the foundation of primate archaeology | Aos ombros de gigantes: O contributo de Cláudia Sousa para a fundação da arqueologia dos primatas". Etnografica 20 3 (2016): 648-652. http://www.scopus.com/inward/record.url?eid=2-s2.0-85003597883&partnerID=MN8TOARS. 10.4000/etnografica.4724 Carvalho, S., Valera, A., Bicho, N., ; Costa, Cláudia; Gonçalves, Célia Gonçalves; Cascalheira, João; Marreiros, João; Pereira, Telmo; Carvalho, Susana; Valera, António; Bicho, Nuno. "ICArEHB - Centro Interdisciplinar de Arqueologia e Evolução do Comportamento Humano um novo polo de investigação arqueológica". Al-Madam 20 2 (2016): 98-101. Published Assis, Sandra; Campanacho, Vanessa; Carvalho, Vânia; Cruz, Cristina Barroso; Curate, Francisco; Gonçalves, David; Marques, Richard; et al. "Looking back at a decade of science communication in the field of human evolution". (2016): http://hdl.handle.net/10316/80397. Hockings KJ; Bryson-Morrison N; Carvalho S; Fujisawa M; Humle T; McGrew WC; Nakamura M; et al. "Tools to tipple: ethanol ingestion by wild chimpanzees using leaf-sponges.". (2015): http://europepmc.org/abstract/med/26543588. 10.1098/rsos.150150 Hockings KJ; McLennan MR; Carvalho S; Ancrenaz M; Bobe R; Byrne RW; Dunbar RI; et al. "Apes in the Anthropocene: flexibility and survival.". (2015): http://europepmc.org/abstract/med/25766059. 10.1016/j.tree.2015.02.002 Benito-Calvo A; Carvalho S; Arroyo A; Matsuzawa T; de la Torre I. "First GIS analysis of modern stone tools used by wild chimpanzees (Pan troglodytes verus) in Bossou, Guinea, West Africa.". (2015): http://europepmc.org/abstract/med/25793642. 10.1371/journal.pone.0121613 Caruana MV; Carvalho S; Braun DR; Presnyakova D; Haslam M; Archer W; Bobe R; Harris JW. "Quantifying traces of tool use: a novel morphometric analysis of damage patterns on percussive tools.". (2014): http://europepmc.org/abstract/med/25415303. 10.1371/journal.pone.0113856 Haslam, M.; Gumert, M.D.; Biro, D.; Carvalho, S.; Malaivijitnond, S.. "Use-Wear Patterns on Wild Macaque Stone Tools Reveal Their Behavioural History". PLoS ONE 8 8 (2013): http://www.scopus.com/inward/record.url?eid=2-s2.0-84881563676&partnerID=MN8TOARS. 10.1371/journal.pone.0072872 Assis, Sandra; Carvalho, Susana. "Evolutionism and the Teaching of Science: How Portugal Has Been “Playing with the Big Tree of Evolution”". Evolution: Education and Outreach 5 3 (2012): 445-452. http://dx.doi.org/10.1007/s12052-012-0437-7. 10.1007/s12052-012-0437-7 Hockings, K.J.; Humle, T.; Carvalho, S.; Matsuzawa, T.. "Chimpanzee interactions with nonhuman species in an anthropogenic habitat". Behaviour 149 3-4 (2012): 299-324. http://www.scopus.com/inward/record.url?eid=2-s2.0-84861675783&partnerID=MN8TOARS. 10.1163/156853912X636735 Carvalho, S.; Biro, D.; Cunha, E.; Hockings, K.; McGrew, W.C.; Richmond, B.G.; Matsuzawa, T.. "Chimpanzee carrying behaviour and the origins of human bipedality". Current Biology 22 6 (2012): http://www.scopus.com/inward/record.url?eid=2-s2.0-84858674149&partnerID=MN8TOARS. 10.1016/j.cub.2012.01.052 Carvalho, Susana; Yamanashi T.; Yamakoshi, G.; Matsuzawa, T.. "Bird in the hand: Bossou chimpanzees capture West African wood-owls (Ciccaba woodfordi) but not to eat.". Pan African News 17 1 (2010): 6-9. Published Haslam, M.; Hernandez-Aguilar, A.; Ling, V.; Carvalho, S.; De La Torre, I.; Destefano, A.; Du, A.; et al. "Primate archaeology". Nature 460 7253 (2009): 339-344. http://www.scopus.com/inward/record.url?eid=2-s2.0-67650796399&partnerID=MN8TOARS. 10.1038/nature08188 Carvalho, S.; Biro, D.; McGrew, W.C.; Matsuzawa, T.. "Tool-composite reuse in wild chimpanzees (Pan troglodytes): Archaeologically invisible steps in the technological evolution of early hominins?". Animal Cognition 12 1 SUPPL (2009): http://www.scopus.com/inward/record.url?eid=2-s2.0-70349776894&partnerID=MN8TOARS. 10.1007/s10071-009-0271-7 Ling, V.; Hernandez-Aguilar, A.; Haslam, M.; Carvalho, S.. "The origins of percussive technology: A smashing time in Cambridge". Evolutionary Anthropology 18 2 (2009): 48-49. http://www.scopus.com/inward/record.url?eid=2-s2.0-68049112468&partnerID=MN8TOARS. 10.1002/evan.20197 Carvalho, S.; Cunha, E.; Sousa, C.; Matsuzawa, T.. "Chaînes opératoires and resource-exploitation strategies in chimpanzee (Pan troglodytes) nut cracking". Journal of Human Evolution 55 1 (2008): 148-163. http://www.scopus.com/inward/record.url?eid=2-s2.0-44749089713&partnerID=MN8TOARS. 10.1016/j.jhevol.2008.02.005 Carvalho, Susana; Sousa, Claudia; Matsuzawa, T.. "New nut-cracking sites in Diecké forest, Guinea: An overview of the etho-archaeological surveys.". Pan African News 14 1 (2007): 11-13. https://ora.ox.ac.uk/objects/uuid:0561b123-daf3-481a-a521-7c10f73038cd/files/m7343d55563da7a17378d4b9f1bcb1898. Open access • Published
Magazine article	Carvalho, Susana. "Anonymous Peer Review: Truth or Trolling?", Scientific American, 2020, https://www.scientificamerican.com/article/anonymous-peer-review-truth-or-trolling/#:~:text=Because%20reviewing%20is%20a%20reciprocal,reviewer%20from%20retaliation%20and%20discrimination.. René Bobe; Carvalho, Susana. "Are humans off the hook for megaherbivore extinctions? Not quite.", The Conversation, 2019, https://theconversation.com/humans-are-not-off-the-hook- for-extinctions-of-large-herbivores-then-or-now-107727. Hockings, Kimberley; Carvalho, Susana. "Apes under pressure show their ingenuity – and hint at our own evolutionary past.", The Conversation, 2015, https://theconversation.com/apes- under-pressure-show-their-ingenuity-and-hint-at-our-own-evolutionary-past-39897.
Online resource	Assis, Sandra; Lopes, Célia; Gonçalves, David; Carvalho, Susana. "Journey to the past/ Viagem ao passado". Actividades pedagógicas dirigidas a crianças e jovens em idade escolar. 2017. https://ln.sync.com/dl/d492eb2b0/fw342igh-apkdi48b-ujferdxs-2czxe8z3. Assis, Sandra; Lopes, Célia; Gonçalves, David; Carvalho, Susana. "Can you identify the bones of my skeleton?/ consegues identificar os meus ossos?" Actividades pedagógicas dirigidas a crianças e jovens em idade escolar.. 2017. https://ln.sync.com/dl/a42b1f550/eeex362i-87ynpryz-6pf25rsq-tswdstce. Assis, Sandra; Lopes, Célia; Gonçalves, David; Carvalho, Susana. "The planet of primates/ planeta dos primatas". Actividades pedagógicas dirigidas a crianças e jovens em idade escolar. 2017. https://ln.sync.com/dl/7666fa970/ju6gdi3h-3m2pydqz-v8qvaic3-rj8rvtw4.
Preprint	Elliot Howard-Spink; Tetsuro Matsuzawa; Susana Carvalho; Catherine Hobaiter; Katarina Almeida-Warren; Thibaud Gruber; Dora Biro. "Old age variably impacts chimpanzee engagement and efficiency in stone tool use". 2025. https://doi.org/10.7554/eLife.105411.1. 10.7554/eLife.105411.1 E Howard-Spink; T Matsuzawa; S Carvalho; C Hobaiter; K Almeida-Warren; T Gruber; D Biro; et al. "Old age variably impacts chimpanzee engagement and efficiency in stone tool use". 2024. https://doi.org/10.1101/2024.11.25.625128. 10.1101/2024.11.25.625128 Sophie Berdugo; Emma Cohen; Arran J. Davis; Tetsuro Matsuzawa; Susana Carvalho. "The ontogeny of chimpanzee technological efficiency". 2024. https://doi.org/10.1101/2024.07.31.605100. 10.1101/2024.07.31.605100 Silva, Maria J Ferreira da; Tralma, Paula; Colmonero-Costeira, Ivo; Mota, Mariana; Farassi, Rassina; Hammond, Philippa; Lewis-Bevan, Lynn; et al. "Sex-mediated gene flow of grayfoot chacma baboons (Papio ursinus griseipes) in a highly seasonal habitat in Gorongosa National Park". 2024. https://doi.org/10.21203/rs.3.rs-4616083/v1. 10.21203/rs.3.rs-4616083/v1 S. Berdugo; E. Cohen; A. J. Davis; T. Matsuzawa; S. Carvalho. "Stable long-term individual variation in chimpanzee technological efficiency". 2023. https://doi.org/10.1101/2023.11.21.568000. 10.1101/2023.11.21.568000 Jana Muschinski; Alexander Mielke; Susana Carvalho. "A network-based analysis of signal use during approach interactions across sexes in chacma baboons (Papio ursinus griseipes)". 2023. https://doi.org/10.1101/2023.02.04.527103. 10.1101/2023.02.04.527103 Alexander Mielke; Susana Carvalho. "Chimpanzee play sequences are structured hierarchically as games". 2022. https://doi.org/10.1101/2022.06.14.496075. 10.1101/2022.06.14.496075 Katarina Almeida-Warren; Tetsuro Matsuzawa; Susana Carvalho. "Landscaping the behavioural ecology of primate stone tool use". 2021. https://doi.org/10.1101/2021.08.24.457369. 10.1101/2021.08.24.457369

Award

2024	BBVA Biodiversity Conservation World Award (to the Gorongosa Restoration Project team - Carvalho is staff on the project) Fundación BBVA, Spain
2021	British Academic Mid-Career Fellowship The British Academy, United Kingdom
2016	Philip Leverhulme Prize Leverhulme Trust, United Kingdom
2014	Honourable Mention for Outstanding Accomplishment in Research The George Washington University, United States
2012	Phyllis and Eileen Gibbs Travelling Fellowship University of Cambridge Newnham College, United Kingdom
2010	Best Portuguese Project in Museology (Team Award)

Title

2021	Professor of Paleoanthropology University of Oxford Institute of Cognitive and Evolutionary Anthropology, United Kingdom

Other distinction

2014	Global Explorer National Geographic Society, United States