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I am a Biologist with a PhD in Molecular and Environmental Biology (2020), conducted at University of California, Davis (USA; research host) and University of Minho (Portugal; degree host), mentored by Prof. Min Zhao. My dissertation focused on the redox–bioelectric interplay during Xenopus regeneration, where I demonstrated that atmospheric oxygen influx generates hydrogen peroxide, which subsequently modulates electrical currents (Development 2016, Nature Communications 2018, Nature Protocols 2020). In parallel, a collaborative effort uncovered the differential electrotaxis of enterobacteria in the mouse gut (Nature Microbiology 2024). Additionally, I contributed to seven other studies exploring the roles of redox and bioelectric cues in the mammalian cornea and rodent gut. As a postdoctoral researcher, I was awarded a long-term European Molecular Biology Organization (EMBO) fellowship to investigate the mechanobiology of Xenopus regeneration (manuscript in preparation) at the Gulbenkian Institute for Molecular Medicine (Portugal). Another research line uncovered a mechanoelectrical mechanism that drives collective cell migration via electrotaxis during Xenopus development (Nature Materials 2025). My research in the Biological Sciences and Biophysics aims to decipher the biochemical–biophysical crosstalk that governs vertebrate regeneration and development. I have applied advanced techniques to study the redox, electrical, and mechanical properties of living organisms. My toolkit includes self-referencing vibrating voltage probes and ion-selective microelectrodes, glass microelectrodes, direct current electric fields, micro-optrodes, microfluidics, laser ablation, tension probes, and in vivo atomic force microscopy. This interdisciplinary approach, combining different biophysical techniques with state of the art molecular biology tools, is central to my work. In the early stages of my career, I have collaborated with research institutions in the USA, Portugal, and Germany, resulting in 12 publications in international peer-reviewed journals (h-index 6, Scopus 2025), 2 preprints, and 3 research funding awards. I have also served as a peer reviewer for academic papers and evaluated a Human Frontier Science Program (HFSP) research grant.
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Identification

Personal identification

Full name
Fernando Ricardo da Silva Ferreira

Citation names

  • Ferreira, Fernando

Author identifiers

Ciência ID
A219-C9BF-5BDB
ORCID iD
0000-0001-5786-8896
Google Scholar ID
https://scholar.google.com/citations?user=Ar2Oo44AAAAJ&hl=pt-PT
Scopus Author Id
56828767000

Addresses

  • Rua da Quinta Grande, 6, 2780-156, Oeiras, Oeiras, Portugal (Professional)

Websites

Knowledge fields

  • Natural sciences - Biological Sciences
  • Natural sciences - Biological Sciences - Biophysics

Languages

Language Speaking Reading Writing Listening Peer-review
Portuguese (Mother tongue)
English Proficiency (C2) Proficiency (C2) Proficiency (C2) Proficiency (C2) Proficiency (C2)
Education
Degree Classification
2011/10/01 - 2020/07/07
Concluded
 Molecular and Environmental Biology (Doutoramento)
University of California Davis Department of Dermatology, United States

Universidade do Minho Centro de Biologia Molecular e Ambiental, Portugal
"Unveiling the integration of redox and bioelectric phenomena in vertebrate regeneration." (THESIS/DISSERTATION)
 Very Good (Best possible)
2014/03/30 - 2014/06/04
Attended
 Chemical Biology of Oxidants and Free Radicals (Outros)
University of California Davis School of Medicine, United States
 Ungraded (attendance-only)
2008/10/01 - 2011/02/28
Concluded
 Evolutionary and Developmental Biology (Mestrado)
Universidade de Lisboa Faculdade de Ciências, Portugal

Gulbenkian Institute for Molecular Medicine, Portugal
"Regeneração da barbatana caudal em peixe-zebra (Danio rerio): uma perspectiva biofísica (Caudal fin regeneration in zebrafish (Danio rerio): a biophysical approach)" (THESIS/DISSERTATION)
 17
2005/10/01 - 2008/06/30
Concluded
 Applied Biology (Licenciatura)
Universidade do Minho Escola de Ciências, Portugal
"Regeneração de extremidades, uma perspectiva da genética e da bioquímica celular - o estado da arte (Appendage regeneration, a genetics and cellular biochemistry approach – the state of the art)" (THESIS/DISSERTATION)
 16,37
Affiliation

Science

Category
Host institution 		Employer
2023/11/01 - Current Postdoc (Research) Gulbenkian Institute for Molecular Medicine, Portugal
Gulbenkian Institute for Molecular Medicine, Portugal
2023/02/16 - 2023/10/31 Postdoc (Research) Universidade Nova de Lisboa Instituto de Tecnologia Química e Biológica António Xavier, Portugal
Universidade Nova de Lisboa Instituto de Tecnologia Química e Biológica António Xavier, Portugal
2020/09/01 - 2023/02/15 Postdoc (Research) Instituto Gulbenkian de Ciência, Portugal
Instituto Gulbenkian de Ciência, Portugal
2019/10/01 - 2020/08/31 Research Assistant (Research) Instituto Gulbenkian de Ciência, Portugal
2011/10/01 - 2020/07/07 Research Trainee (Research) Universidade do Minho Centro de Biologia Molecular e Ambiental, Portugal
Universidade do Minho Centro de Biologia Molecular e Ambiental, Portugal
2017/09/11 - 2018/04/30 Research Assistant (Research) University of California Davis Department of Dermatology, United States
University of California Davis Department of Dermatology, United States
2013/06/01 - 2017/09/10 Research Trainee (Research) University of California Davis Department of Dermatology, United States
University of California Davis Department of Dermatology, United States
2014/06/12 - 2014/08/02 Visiting Researcher (Research) Marine Biological Laboratory, United States
Marine Biological Laboratory, United States
2009/06/15 - 2010/10/29 Research Trainee (Research) Gulbenkian Institute for Molecular Medicine, Portugal
Gulbenkian Institute for Molecular Medicine, Portugal
Projects

Grant

Designation Funders
2020/09/01 - 2022/10/31 Mechanical control of collective cell motion in vertebrate regeneration
EMBO ALTF 27-2020
Gulbenkian Institute for Molecular Medicine, Portugal
 European Molecular Biology Laboratory
Concluded
2020/08 - 2020/08 Unveiling the mechanical nature of tissue regeneration
E200300401
European Soft Matter Infrastructure, Germany
 European Soft Matter Infrastructure
Concluded
2013/06/01 - 2017/05/31 Tail regeneration in Xenopus laevis tadpoles: A bioelectrical and oxidative approach
35256DFA
SFRH/BD/87256/2012
University of California Davis Department of Dermatology, United States
 Fundação para a Ciência e a Tecnologia
Concluded
Outputs

Publications

Conference paper
  1. Li Ma; Fernando Ferreira; Brian Reid; Min Zhao. "An oxygen-specific optical “electrode” (optrode) directly measures oxygen uptake and reveals its dynamics at ocular surface". 2017.
  2. Y. Shen; T. Pfluger; Fernando Ferreira; J. Liang; M. Navedo; Q. Zeng; B. Reid; M. Zhao. "Wound electric signals are significantly reduced in diabetic animal". 2016.
    10.1016/j.jid.2016.05.084
  3. Yaohui Sun; Kan Zhu; Yuxin Sun; Brian Reid; Fernando Ferreira; Y. Li; X. Gao; et al. "Individual and collective cell polarization and migration in electric field". 2015.
    10.1091/mbc.e15-09-0674
Encyclopedia entry
  1. Levin, Michael; Bates, Emily; Zhao, Min; Ferreira, Fernando. "Developmental bioelectricity". In Wikipedia, The Free Encyclopedia, edited by Wikipedia contributors. Wikipedia, The Free Encyclopedia, 2018. https://en.wikipedia.org/wiki/Developmental_bioelectricity.
    Published
Journal article
  1. Fernando Ferreira; Sofia Moreira; Min Zhao; Elias H. Barriga. "Stretch-induced endogenous electric fields drive directed collective cell migration in vivo". Nature Materials (2025): https://doi.org/10.1038/s41563-024-02060-2.
    10.1038/s41563-024-02060-2
  2. Li Ma; Fernando Ferreira; Brian Reid; Liang Guo; Min Zhao. "Optical microsensing reveals spatiotemporal oxygen dynamics in cornea wounds that affect healing via reactive oxygen species". The FASEB Journal (2024): https://doi.org/10.1096/fj.202401054R.
    10.1096/fj.202401054R
  3. Yaohui Sun; Fernando Ferreira; Brian Reid; Kan Zhu; Li Ma; Briana M. Young; Catherine E. Hagan; et al. "Gut epithelial electrical cues drive differential localization of enterobacteria". Nature Microbiology (2024): https://doi.org/10.1038/s41564-024-01778-8.
    10.1038/s41564-024-01778-8
  4. Sun Qin; Li Ma; Fernando Ferreira; Chelsea Brown; Manuel F. Navedo; Brian Reid; Min Zhao. "Diabetic Ocular Surface Has Defects in Oxygen Uptake Revealed by Optic Fiber Microsensor". Investigative Opthalmology & Visual Science 65 3 (2024): 27-27. http://dx.doi.org/10.1167/iovs.65.3.27.
    10.1167/iovs.65.3.27
  5. Yaohui Sun; Brian Reid; Yan Zhang; Kan Zhu; Fernando Ferreira; Alejandro Estrada; Yuxin Sun; et al. "Electric field-guided collective motility initiation of large epidermal cell groups". Molecular Biology of the Cell (2023): https://doi.org/10.1091/mbc.E22-09-0391.
    10.1091/mbc.E22-09-0391
  6. Qin Sun; Li Ma; Fernando Ferreira; Chelsea Brown; Brian Reid; Min Zhao. "Optic Fiber Microsensor Reveals Specific Spatiotemporal Oxygen Uptake Profiles at the Mammalian Ocular Surface". Biosensors (2023): https://doi.org/10.3390/bios13020245.
    10.3390/bios13020245
  7. Fernando Ferreira; Guillaume Luxardi; Brian Reid; Li Ma; VijayKrishna Raghunathan; Min Zhao. "Real-time physiological measurements of oxygen using a non-invasive self-referencing optical fiber microsensor". Nature Protocols (2020): https://doi.org/10.1038/s41596-019-0231-x.
    10.1038/s41596-019-0231-x
  8. Yaohui Sun; David S. Schneider; Brian Reid; Fernando Ferreira; Guillaume Luxardi; Li Ma; Kristen L. Lokken; et al. "Infection-generated electric field in gut epithelium drives bidirectional migration of macrophages". PLOS Biology 17 4 (2019): e3000044-e3000044. https://doi.org/10.1371/journal.pbio.3000044.
    10.1371/journal.pbio.3000044
  9. Fernando Ferreira; VijayKrishna Raghunathan; Guillaume Luxardi; Kan Zhu; Min Zhao. "Early redox activities modulate Xenopus tail regeneration". Nature Communications 9 1 (2018): https://doi.org/10.1038/s41467-018-06614-2.
    10.1038/s41467-018-06614-2
  10. Fernando Ferreira; Guillaume Luxardi; Brian Reid; Min Zhao. "Early bioelectric activities mediate redox-modulated regeneration". Development (2016): https://doi.org/10.1242/dev.142034.
    10.1242/dev.142034
  11. Yunyun Shen; Trisha Pfluger; Fernando Ferreira; Jiebing Liang; Manuel F. Navedo; Qunli Zeng; Brian Reid; Min Zhao. "Diabetic cornea wounds produce significantly weaker electric signals that may contribute to impaired healing". Scientific Reports (2016): https://doi.org/10.1038/srep26525.
    10.1038/srep26525
  12. Guillaume Luxardi; Brian Reid; Fernando Ferreira; Pauline Maillard; Min Zhao. "Measurement of Extracellular Ion Fluxes Using the Ion-selective Self-referencing Microelectrode Technique". Journal of Visualized Experiments (2015): https://dx.doi.org/10.3791/52782-v.
    10.3791/52782
Preprint
  1. Ferreira, Fernando; Moreira, Sofia; Zhao, M.; Barriga, Elias H.. "Stretch-induced endogenous electric fields drive neural crest directed collective cell migration in vivo". 2021. http://dx.doi.org/10.1101/2021.10.11.463916.
    10.1101/2021.10.11.463916
  2. Yaohui Sun; Brian Reid; Fernando Ferreira; Guillaume Luxardi; Li Ma; Kristen L. Lokken; Kan Zhu; et al. "Infection-generated electric field in gut epithelium drives bidirectional migration of macrophages". 2018. https://doi.org/10.1101/427294.
    10.1101/427294
Thesis / Dissertation
  1. "Unveiling the integration of redox and bioelectric phenomena in vertebrate regeneration". PhD, 2020. https://hdl.handle.net/1822/75988.
  2. Ferreira, Fernando Ricardo da Silva, 1987-. "Regeneração da barbatana caudal em peixe-zebra (Danio rerio): uma perspectiva biofísica". Master, Universidade de Lisboa Departamento de Biologia Animal, 2011. http://hdl.handle.net/10451/2918.

Other

Software
  1. Ferreira, Fernando; Eugénia Fernandes; José Correia; Susana Silva. "App Jardim • Vertigem (ir)reversível (Garden • (Ir)reversible Vertigo)". v1.0. Google Play Android. 2019.
Activities

Oral presentation

Presentation title Event name
Host (Event location)
2024/06/19 Mechano-molecular synchronisation of Xenopus laevis tail regeneration EMBO workshop – Limb Development: Fundamental mechanisms, evolution, disease and regeneration
European Molecular Biology Organization (Dresden, Germany)
2021/05/19 Steering cells electrically: the spark that guides neural crest migration Postdoc Committee Seminar
Gulbenkian Institute of Science (Oeiras, Portugal)
2020/07/29 Mechanical control of regeneration Undergraduate IGC Summer School 2020
Gulbenkian Institute of Science (Oeiras, Portugal)
2019/07/26 An early crosstalk on regeneration: bioelectric and redox signalling 78th Annual Society for Developmental Biology Meeting; SDB Satellite Symposium on Developmental Bioelectricity
Society for Developmental Biology (Boston, United States)
2017/11/15 Redox and bioelectric probing: harnessing the sensing power of photons and electrons UC Davis School of Medicine Department of Dermatology Research Symposium 2017
University of California, Davis Department of Dermatology (Sacramento, United States)
2016/06/22 Early HIF-1a stabilization, potentially ROS-dependent, modulates regeneration UC Davis School of Medicine Dermatology Symposium 2016
University of California, Davis Department of Dermatology (Sacramento, United States)
2016/05/25 Early bioelectric activities mediate redox-modulated regeneration UC Davis Department of Dermatology EpiBio Meeting
University of California, Davis Department of Dermatology (Sacramento, United States)

Supervision

Thesis Title
Role 		Degree Subject (Type)
Institution / Organization
2020/09 - 2022/01 Mechanical synchronization of Xenopus tail regeneration (http://hdl.handle.net/10451/51898)
Co-supervisor
 Evolutionary and Developmental Biology (Master)
Universidade de Lisboa Departamento de Biologia Animal, Portugal

Gulbenkian Institute for Molecular Medicine, Portugal

Event organisation

Event name
Type of event (Role) 		Institution / Organization
2008 - 2008 XXII ENEB Encontro Nacional de Estudantes de Biologia (National Meeting of Portuguese Biology Students) (2008 - 2008)
Conference (Member of the Scientific Committee)
 Universidade do Minho Escola de Ciências, Portugal

Event participation

Activity description
Type of event 		Event name
Institution / Organization
2024/05/16 - 2024/05/16 ERC Grant Writing Course (5 sessions, 6 h) instructed by Randy Levinson, PhD (Lai Wen Di Consulting)
Workshop
 ERC Grant Writing Course
LS4FUTURE, Portugal
2024/04/15 - 2024/04/18 EMBL Symposium: The mechanics of life: from development to disease (Virtual)
Symposium
 EMBL Symposium: The mechanics of life: from development to disease
European Molecular Biology Organization (EMBO), Germany

European Molecular Biology Laboratory, Germany
2023/07/03 - 2023/07/04 EMBO Fellows’ Meeting 2023 (Heidelberg, Germany)
Meeting
 EMBO Fellows’ Meeting 2023
European Molecular Biology Organization (EMBO), Germany
2023/07/03 - 2023/07/04 Development and Morphogenesis: Symposium in honour of Maria Leptin (Heidelberg, Germany)
Symposium
 Development and Morphogenesis: Symposium in honour of Maria Leptin
European Molecular Biology Organization (EMBO), Germany
2022/09/22 - 2022/11/24 Webinar Series How to streamline the crafting of a scientific paper (10 lectures, 15 h, online) instructed by Márcia Triunfal, PhD (Publicase International)
Workshop
 Webinar Series How to Streamline the Crafting of a Scientific Paper
Publicase International, Portugal
2022/11/17 - 2022/11/17 Cybersecurity awareness seminar (1 session, 1 h, online) instructed by Member of S21SEC Academy
Seminar
 Cybersecurity awareness
Fundação Calouste Gulbenkian, Portugal
2022/11/15 - 2022/11/16 EMBO Solutions Course How to review a scientific paper (2 sessions, 8 h, online) instructed by Lise Roth, PhD (Senior Scientific Editor EMBO Molecular Medicine)
Workshop
 EMBO How to review a scientific paper
European Molecular Biology Organization (EMBO), Germany
2022/10/16 - 2022/10/20 Poster presentation (selected by jury) at the 19th International Congress of Developmental Biology (ISDB) (Algarve, Portugal)
Congress
 Stretch-induced endogenous electric fields drive directed collective cell migration in vivo
International Society of Developmental Biologists, United States
2022/09/29 - 2022/09/30 Poster presentation (selected by jury) at the 16th Physics Of Living Matter Symposium (Cambridge, United Kingdom)
Symposium
 Mechanically induced electric fields drive directed collective cell migration in vivo
University of Cambridge Cambridge Centre for Physical Biology, United Kingdom
2022/09/27 - 2022/09/28 Poster presentation (selected by jury) at the 16th Physics Of Living Matter Symposium (Virtual from Marseille)
Symposium
 Mechanically induced electric fields drive directed collective cell migration in vivo
Aix-Marseille Université, France
2022/05/25 - 2022/05/27 Poster presentation at the Scientific Advisory Board Visit (Oeiras, Portugal)
Other
 Mechanisms of Morphogenesis Laboratory: Projects and Tools
Gulbenkian Institute for Molecular Medicine, Portugal
2022/02/01 - 2022/02/01 General data protection regulation (1 session, 1 h, online) instructed by lawyer from Viera de Almeida firm
Seminar
 General data protection regulation
Fundação Calouste Gulbenkian, Portugal
2021/06/22 - 2021/06/24 EMBO Solutions Course EMBO Laboratory Leadership (3 sessions, 22 h, online) instructed by Melanie Glickman and Alexandra Hund
Workshop
 EMBO Laboratory Leadership
European Molecular Biology Organization (EMBO), Germany
2020/09/15 - 2020/09/15 Research Integrity – Natural and Physical Sciences course (1 session, 6 h, online) using web-based platform (Epigeum Online Course System)
Workshop
 Research Integrity – Natural and Physical Sciences
Oxford University Press, United Kingdom

Epigeum, United Kingdom
2020/07/27 - 2020/07/31 Science outreach
Other
 Undergraduate IGC Summer School 2020
Gulbenkian Institute for Molecular Medicine, Portugal
2019/10/07 - 2019/10/17 SPCAL Laboratory Animal Sciences (LAS) in Mouse, Zebrafish and Xenopus (Category B and Functions A + D) (20 theoretical lectures, 20 h, online and 6 practical sessions at Gulbenkian Institute of Science) instructed by Manuel Rebelo, PhD; Ana Sofia Leocádio, BSc; Joana Bom, MSc; Cristina Borges, PhD; Maysa Franco, MSc; Inês Santos, MSc; Liliana Vale, BSc; and Nuno Pereira, MV
Workshop
 SPCAL Laboratory Animal Sciences (LAS) in Mouse, Zebrafish and Xenopus
Congento Consortium for Genetically Tractable Organisms, Portugal

Gulbenkian Institute for Molecular Medicine, Portugal
2017/12/21 - 2017/12/21 Poster presentation at the A decade integrating biology (in) research and training: 10 years of the MSc in Evolutionary and Developmental Biology (BED) (Lisbon, Portugal)
Symposium
 Integration of early redox and bioelectric phenomena modulates vertebrate regeneration
Universidade de Lisboa Departamento de Biologia Animal, Portugal
2017/11/15 - 2017/11/15 Poster presentation at the UC Davis School of Medicine Department of Dermatology Research Symposium 2017 (Sacramento, California, United States)
Symposium
 Redox and bioelectric probing: harnessing the sensing power of photons and electrons
University of California Davis Department of Dermatology, United States
2016/06/22 - 2016/06/22 Poster presentation at the UC Davis School of Medicine Dermatology Symposium 2016 (Sacramento, California, United States)
Symposium
 Early HIF-1a stabilization, potentially ROS-dependent, modulates regeneration
University of California Davis Department of Dermatology, United States
2016/05/04 - 2016/05/06 Poster presentation (selected by jury) at the Oxidants and Antioxidants in Biology: Redox Medicine and Nutrition (Davis, California, United States)
Conference
 Reactive oxygen species regulate bioelectric responses during tail regeneration of Xenopus laevis tadpoles
Oxygen Club of California, United States
2014/12/10 - 2014/12/10 Poster presentation at the UC Davis School of Medicine Dermatology Symposium 2014 (Sacramento, California, United States)
Symposium
 Reactive oxygen species regulate bioelectric responses in tail regeneration of Xenopus laevis tadpoles
University of California Davis Department of Dermatology, United States
2014/07/06 - 2014/07/11 Poster presentation (selected by jury) at the Bioelectrochemistry: Cellular and Organismal Responses to Endogenous and Exogenous Fields (Biddeford, Maine, United States)
Conference
 Mapping bioelectric phenomena during tail regeneration in Xenopus laevis tadpoles
Gordon Research Conference, United States
2010/10 - 2010/10 Poster presentation (selected by jury) at the Joint Meeting of the Portuguese and Spanish Developmental Biology Societies (JMPSDBS) (Badajoz, Spain)
Meeting
 Extracellular chloride and sodium flux dynamics during fin regeneration in zebrafish
Sociedade Portuguesa de Biologia do Desenvolvimento, Portugal

Sociedad Española de Biología del Desarrollo, Spain
2009/10/10 - 2009/10/10 Science outreach
Other
 Open Day 2009
Gulbenkian Institute for Molecular Medicine, Portugal

Ad Hoc journal article review

Journal title (ISSN) Publisher
2020 - Current Frontiers in Cell and Developmental Biology (2296-634X) Frontiers Media
2024 - 2024 Redox Biology (2213-2317) Elsevier

Consulting

Activity description Institution / Organization
2018 - 2018 Critical reading and editing (one chapter's section in Ahead of the Curve, Volume 2 Hidden breakthroughs in the biosciences) IOPscience, United Kingdom
2018 - 2018 Reviewer assistance (Technique manuscript) (IEEE Access) IEEE, United States
2017 - 2017 Reviewer assistance (Research manuscript) (Biophysical Journal) Cell Press, United States

Course / Discipline taught

Academic session Degree Subject (Type) Institution / Organization
2021/01/15 - 2021/01/15 Lecture in the form of a ~1 h seminar for undergraduate students in the course "Differentition and Development" (Portuguese: "Diferenciação e Desenvolvimento") (invited by the course instructor) Biologia Aplicada (Bacharelato) Universidade do Minho Escola de Ciências, Portugal

Evaluation committee

Activity description
Role 		Institution / Organization Funding entity
2023 - 2023 Reviewer (2024 HFSP Research Grant Awards)
Evaluator
 Human Frontier Science Program, France Human Frontier Science Program

Interview (newspaper / magazine)

Activity description Newspaper / Forum
2025/01/17 Brief interview for the press release 'Shocking Cues: How Cells Harness Electric Fields to Migrate During Embryonic Development' on the publication "Stretch-induced endogenous electric fields drive directed collective cell migration in vivo. Nature Materials. 2025" PoL Dresden University of Technology News portal
2018/11/07 Brief interview for the press release 'Exploring the Role of Redox and Bioelectric Players in Tissue Regeneration' on the publication "Early redox activities modulate Xenopus tail regeneration. Nature Communications. 2018" UC Davis Egghead blog
2016/12/14 Interview for the series "The people behind the papers" on the publication "Early bioelectric activities mediate redox-modulated regeneration. Development. 2016" The Node Interviews

Mentoring / Tutoring

Topic Student name
2020/01 - 2020/06 Mechanosensitive channels on Xenopus laevis tadpole tail regeneration Quentin Tirel
2016/02 - 2016/12 Bioelectricity on Xenopus laevis tadpole tail regeneration Garrett Chan
2015/05 - 2016/05 Ion channels on Xenopus laevis wound healing Zachary Chak
Distinctions

Award

2020 EMBO Long-term Postdoctoral fellowship
European Molecular Biology Organization (EMBO), Germany
2020 EUSMI Transnational Access Programme
European Soft Matter Infrastructure (EUSMI), Germany
2019 Favourite Garden
International Garden Festival of Ponte de Lima, Portugal
2012 PhD Fellowship
Fundação para a Ciência e a Tecnologia, Portugal