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Eliza Fragkopoulou is a postdoc researcher at CCMAR, University of the Algarve, studying how climate change is affecting and will further affect the distribution and genetic diversity of marine species, at global scales. Her work is focused on marine species distribution models for past, present and future conditions based on different climate change scenarios. She has experience in meta-analyses and biodiversity data compilation under the framework of open science and the FAIR principle for findable, accessible, interoperable, and reusable data. The outcomes of her research are completely aligned with the 2030 Agenda for Sustainable Development, adopted by all United Nations Members. Working at the Center of Marine Sciences (recurrently classified as excellent in national evaluations) has resulted in collaborations with several national and international teams (e.g., from Norway, Brazil, Saudi Arabia, Australia and Denmark).
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Identificação

Identificação pessoal

Nome completo
Eliza Fragkopoulou

Nomes de citação

  • Fragkopoulou, Eliza

Identificadores de autor

Ciência ID
5B1F-9922-2CB5
ORCID iD
0000-0002-0557-3954

Endereços de correio eletrónico

  • eli_frag@hotmail.com (Profissional)

Websites

Domínios de atuação

  • Ciências Naturais - Ciências da Terra e do Ambiente - Oceanografia

Idiomas

Idioma Conversação Leitura Escrita Compreensão Peer-review
Grego Utilizador proficiente (C2) Utilizador proficiente (C2) Utilizador proficiente (C2) Utilizador proficiente (C2)
Inglês Utilizador proficiente (C2) Utilizador proficiente (C2) Utilizador proficiente (C2) Utilizador proficiente (C2)
Português Utilizador proficiente (C2) Utilizador proficiente (C2) Utilizador proficiente (C2) Utilizador proficiente (C2)
Sérvio Utilizador independente (B1) Utilizador independente (B1) Utilizador independente (B1) Utilizador independente (B1)
Francês Utilizador elementar (A2) Utilizador elementar (A2) Utilizador elementar (A2) Utilizador elementar (A2)
Formação
Grau Classificação
2020 - 2024
Concluído
 Ciências do Mar, da Terra e do Ambiente (Doutoramento)
Universidade do Algarve Faculdade de Ciências e Tecnologia, Portugal
"Potential consequences of future climate changes on the marine forest gene pools" (TESE/DISSERTAÇÃO)
 Highest distinction
2013 - 2015
Concluído
 Biologia Marinha (Mestrado)
Especialização em Sem especialização
Universidade do Algarve Faculdade de Ciências e Tecnologia, Portugal
"A GLOBAL ASSESSMENT OF THE EFFECTIVENESS OF DIFFERENT PROTECTION LEVELS IN MARINE PROTECTED AREAS OF TROPICAL WATERS, FOLLOWING A NEW CATEGORIZATION" (TESE/DISSERTAÇÃO)
 17/20 Very Good
2012
Concluído
 Oceanography and Marine Sciences (Bachelor)
University of the Aegean School of Sciences, Grécia
"Exploring the use of satellite images to determine the bathymetry of Kalloni Bay, Lesvos Island." (TESE/DISSERTAÇÃO)
 7.1/10 Very good
Percurso profissional

Ciência

Categoria Profissional
Instituição de acolhimento 		Empregador
2024/05/01 - Atual Pós-doutorado (Investigação) Centro de Ciências do Mar, Portugal
Centro de Ciências do Mar, Portugal

Outros

Categoria Profissional
Instituição de acolhimento 		Empregador
2020/02/01 - 2024/07/18 PhD student Centro de Ciências do Mar, Portugal
Universidade do Algarve - Campus de Gambelas, Portugal
2018/07/15 - 2020/01/31 Research technician under the project MAR-01.04.02- FEAMP- Development of tools for the design of a network of marine protected areas for the integrated management of coastal zones. Universidade do Algarve, Portugal
2015/05/01 - 2018/03/28 Leading Marine Biologist Guide. Organising eco-tours in the Ria Formosa Natural Park, guiding snorkeling/ bird watching groups, demonstrating traditional fishing gear, clam and oyster cultures. Paginas de Verão, Lda, Portugal
2014/11/01 - 2015/01/30 Intern diver/researcher. Sea urchin density monitoring and cartography of a biodiversity hotspot in Ria Formosa lagoon, Portugal. Species identification, density measurements, and samples collection. Universidade do Algarve, Portugal
2014/10/01 - 2014/12/20 Intern researcher. Species distribution modeling of the sea urchin Echinus acutus using basic Maxent. Centro de Ciências do Mar, Portugal
2014/09/01 - 2014/10/30 Intern diver. Seasonal underwater visual census of demersal fish in the Arrabida Marine Park Portugal. Centro de Ciências do Mar, Portugal
2012/11/01 - 2013/02/20 Volunteer. Mapping the high-risk areas for Caretta caretta turtles around Kefalonia Island, Greece. Collecting presence data of injured and/or dead turtles and associating them to main shipping routes Municipality of Kefalonia, Grécia
2012/09/01 - 2012/09/30 Assistant member crew on expedition for mapping the Posidonia oceanica meadows around the South coast of Kefalonia Island, Greece. Use of side scan sonar, sub bottom profiler, and drop of camera. University of Patras, Grécia
2010/05/01 - 2012/02/25 Member of Marine Remote Sensing Group, participating in the expedition for exploring the Kalloni Bay, of Lesvos Island, Greece. Use of side scan sonar, sub bottom profiler and bathymeter. Panepistemio Aigaiou Tmema Okeanographias kai Thalassion Bioepistemon, Grécia
2011/03/01 - 2011/07/01 Outdoor behavioural observer on seals. Monitoring seal activity, count and identification of seal species, recording behavioural patterns and interactions with anglers, monitoring the seals' responses University of Aberdeen Oceanlab, Reino Unido
Projetos

Bolsa

Designação Financiadores
2022 - Atual Restore Seas (EU BiodivRestore-253)
DivRestore/0013/2020
Bolseiro de Doutoramento
Centro de Ciências do Mar, Portugal
Em curso
2021/03 - 2024/02 O Acordo de Paris e a redistribuição da biodiversidade marinha a nível global
Provided by PTCRIS: PTDC/BIA-CBI/6515/2020
 Fundação para a Ciência e a Tecnologia
2020/02 - 2024/01 Potential consequences of future climate changes for marine forest gene pools
Provided by PTCRIS: SFRH/BD/144878/2019
 Fundação para a Ciência e a Tecnologia
2018 - 2020 REDAMP - Development of tools for the design of MPA networks for the integrated management of coastal areas
MAR-01.04.02-FEAMP-0015 REDAMP
 N/A

Projeto

Designação Financiadores
2024/05/01 - Atual iCulture: A digital bio-platform and co-culture bioprocess to prospect and utilise macroalgae responsibly and sustainably
EU Horizon iCULTURE 101082010
Investigador Pós-doutorado
Centro de Ciências do Mar, Portugal
Em curso
2018/07 - 2020/01 REDAMP- Developing tools for the successful design and implementation of coastal networks of Marine Protected Areas (MPAs) in Portugal
MAR-01.04.02-FEAMP-0015
Bolseiro de Técnico de Investigação

Outro

Designação Financiadores
2021 - 2024 PARIS - The Paris Agreement and the global redistribution of marine biodiversity
PTDC/BIA-CBI/6515/2020
Bolseiro de Doutoramento
Concluído
2013 - 2015 BUFFER- Partially protected areas as buffers to increase the linked social-ecological resilience
Biodiversa
Bolseiro de Mestrado
Produções

Publicações

Artigo em revista
  1. Lidiane Gouvêa; Eliza Fragkopoulou; Miguel B. Araújo; Ester A. Serrão; Jorge Assis. "Seagrass Biodiversity Under the Latest-Generation Scenarios of Projected Climate Change". Journal of Biogeography (2024): http://dx.doi.org/10.1111/jbi.15021.
    10.1111/jbi.15021
  2. Jorge Assis; Ester A. Serrão; Eliza Fragkopoulou; Térence Legrand; Lidiane Gouvêa; Miguel B. Araújo. "Misconception of model transferability precludes estimates of seagrass community reorganization in a changing climate". Nature Plants (2024): https://doi.org/10.1038/s41477-024-01735-7.
    10.1038/s41477-024-01735-7
  3. Térence Legrand; Eliza Fragkopoulou; Lauren Vapillon; Lidiane Gouvêa; Ester A. Serrão; Jorge Assis. "Unravelling the role of oceanographic connectivity in the distribution of genetic diversity of marine forests at the global scale". Global Ecology and Biogeography (2024): https://doi.org/10.1111/geb.13857.
    10.1111/geb.13857
  4. João Neiva; Jorge Assis; Eliza Fragkopoulou; Gareth A. Pearson; Peter T. Raimondi; Laura Anderson; Dorte Krause-Jensen; et al. "Trans-Arctic asymmetries, melting pots and weak species cohesion in the low-dispersal amphiboreal seaweed Fucus distichus". Frontiers in Ecology and Evolution (2024): http://dx.doi.org/10.3389/fevo.2024.1356987.
    10.3389/fevo.2024.1356987
  5. Jorge Assis; Eliza Fragkopoulou; Lidiane Gouvêa; Miguel B. Araújo; Ester A. Serrão. "Kelp forest diversity under projected end-of-century climate change". Diversity and Distributions (2024): https://doi.org/10.1111/ddi.13837.
    10.1111/ddi.13837
  6. Ariadni Vafeiadou; Eliza Fragkopoulou; Jorge Assis. "A global dataset of demosponge distribution records". Data in Brief (2024): https://doi.org/10.1016/j.dib.2024.110200.
    10.1016/j.dib.2024.110200
  7. Jorge Assis; Salvador Jesús Fernández Bejarano; Vinícius W. Salazar; Lennert Schepers; Lidiane Gouvêa; Eliza Fragkopoulou; Frederic Leclercq; et al. "Bio-ORACLE v3.0. Pushing marine data layers to the CMIP6 Earth System Models of climate change research". Global Ecology and Biogeography (2024): https://doi.org/10.1111/geb.13813.
    10.1111/geb.13813
  8. Lidiane Gouvêa; Eliza Fragkopoulou; Térence Legrand; Ester A. Serrão; Jorge Assis. "Range map data of marine ecosystem structuring species under global climate change". Data in Brief (2024): https://doi.org/10.1016/j.dib.2023.110023.
    10.1016/j.dib.2023.110023
  9. Eliza Fragkopoulou; Alex Sen Gupta; Mark John Costello; Thomas Wernberg; Miguel B. Araújo; Ester A. Serrão; Olivier De Clerck; Jorge Assis. "Marine biodiversity exposed to prolonged and intense subsurface heatwaves". Nature Climate Change (2023): https://doi.org/10.1038/s41558-023-01790-6.
    10.1038/s41558-023-01790-6
  10. "Intense and prolonged subsurface marine heatwaves pose risk to biodiversity". Nature Climate Change (2023): http://dx.doi.org/10.1038/s41558-023-01796-0.
    10.1038/s41558-023-01796-0
  11. Eliza Fragkopoulou; Alex Sen Gupta; Mark Costello; Thomas Wernberg; Miguel B. Araújo; Ester Serrão; Olivier De Clerck; Jorge Assis. "Marine biodiversity exposed to prolonged and intense subsurface heatwaves". (2023): https://figshare.com/articles/journal_contribution/Marine_biodiversity_exposed_to_prolonged_and_intense_subsurface_heatwaves/19174985.
    10.6084/m9.figshare.19174985
  12. Viktória Balogh; Eliza Fragkopoulou; Ester A. Serrão; Jorge Assis. "A dataset of cold-water coral distribution records". Data in Brief (2023): https://doi.org/10.1016/j.dib.2023.109223.
    10.1016/j.dib.2023.109223
  13. Lidiane P. Gouvêa; Eliza Fragkopoulou; Kyle Cavanaugh; Ester A. Serrão; Miguel B. Araújo; Mark John Costello; E. H. Taraneh Westergerling; Jorge Assis. "Oceanographic connectivity explains the intra-specific diversity of mangrove forests at global scales". Proceedings of the National Academy of Sciences (2023): https://doi.org/10.1073/pnas.2209637120.
    10.1073/pnas.2209637120
  14. David Abecasis; Eliza Fragkopoulou; Bruno Claro; Jorge Assis. "Biophysical modelling and graph theory identify key connectivity hubs in the Mediterranean marine reserve network". Frontiers in Marine Science 9 (2023): http://dx.doi.org/10.3389/fmars.2022.1000687.
    10.3389/fmars.2022.1000687
  15. L. Marquez; E. Fragkopoulou; K. C. Cavanaugh; H. F. Houskeeper; J. Assis. "Artificial intelligence convolutional neural networks map giant kelp forests from satellite imagery". Scientific Reports 12 1 (2022): http://dx.doi.org/10.1038/s41598-022-26439-w.
    10.1038/s41598-022-26439-w
  16. Assis, Jorge; Serrao, Ester; Duarte, Carlos M.; Fragkopoulou, Eliza; Krause-Jensen, Dorte. "Major expansion of marine forests in a warmer Arctic". Frontiers in Marine Science (2022):
    10.3389/fmars.2022.850368
  17. Gouvêa, Lidiane P.; Horta, Paulo A.; Fragkopoulou, Eliza; Gurgel, Carlos F. D.; Peres, Leticia M. C.; Bastos, Eduardo; Ramlov, Fernanda; et al. "Phenotypic Plasticity in Sargassum Forests May Not Counteract Projected Biomass Losses Along a Broad Latitudinal Gradient". Ecosystems (2022): http://dx.doi.org/10.1007/s10021-022-00738-9.
    10.1007/s10021-022-00738-9
  18. Fragkopoulou, Eliza; Serrão, Ester A.; De Clerck, Olivier; Costello, Mark J.; Araújo, Miguel B.; Duarte, Carlos M.; Krause-Jensen, Dorte; Assis, Jorge. Autor correspondente: Serrão, Ester A.. "Global biodiversity patterns of marine forests of brown macroalgae". Global Ecology and Biogeography (2022): http://dx.doi.org/10.1111/geb.13450.
    10.1111/geb.13450
  19. Assis, Jorge; Failler, Pierre; Fragkopoulou, Eliza; Abecasis, David; Touron-Gardic, Gregoire; Regalla, Aissa; Sidina, Ebaye; Dinis, Herculano; Serrao, Ester A.. Autor correspondente: Assis, Jorge. "Potential Biodiversity Connectivity in the Network of Marine Protected Areas in Western Africa". Frontiers in Marine Science 8 (2021): http://dx.doi.org/10.3389/fmars.2021.765053.
    10.3389/fmars.2021.765053
  20. Assis, J.; Fragkopoulou, E.; Serrão, E.A.; Horta e Costa, B.; Gandra, M.; Abecasis, D.. "Weak biodiversity connectivity in the European network of no-take marine protected areas". Science of The Total Environment 773 (2021): 145664. http://dx.doi.org/10.1016/j.scitotenv.2021.145664.
    10.1016/j.scitotenv.2021.145664
  21. Fragkopoulou, Eliza; Serrão, Ester A.; Horta, Paulo A.; Koerich, Gabrielle; Assis, Jorge. Autor correspondente: Fragkopoulou, Eliza. "Bottom Trawling Threatens Future Climate Refugia of Rhodoliths Globally". Frontiers in Marine Science 7 (2021): http://dx.doi.org/10.3389/fmars.2020.594537.
    Acesso aberto • 10.3389/fmars.2020.594537
  22. Assis, Jorge; Fragkopoulou, Eliza; Frade, Duarte; Neiva, João; Oliveira, André; Abecasis, David; Faugeron, Sylvain; Serrão, Ester A.. "A fine-tuned global distribution dataset of marine forests". Scientific Data 7 1 (2020): http://dx.doi.org/10.1038/s41597-020-0459-x.
    Acesso aberto • 10.1038/s41597-020-0459-x
  23. Pilczynska, Joanna; Cocito, Silvia; Boavida, Joana; Serrão, Ester A.; Assis, Jorge; Fragkopoulou, Eliza; Queiroga, Henrique. Autor correspondente: Fragkopoulou, Eliza. "Genetic diversity increases with depth in red gorgonian populations of the Mediterranean Sea and the Atlantic Ocean". PeerJ 7 (2019): e6794. http://dx.doi.org/10.7717/peerj.6794.
    10.7717/peerj.6794
  24. Zupan, Mirta; Fragkopoulou, Eliza; Claudet, Joachim; Erzini, Karim; Horta e Costa, Bárbara; Gonçalves, Emanuel J. "Marine partially protected areas: drivers of ecological effectiveness". Frontiers in Ecology and the Environment 16 7 (2018): 381-387. http://dx.doi.org/10.1002/fee.1934.
    Publicado • 10.1002/fee.1934
Capítulo de livro
  1. Keith, DA; Nelson, W ; Fragkopoulou, Eliza; Assis, Jorge; Schubert, Nadine. "M1.10 Rhodolith/Maërl beds". In IUCN Global Ecosystem Typology 2.0 : descriptive profiles for biomes and ecosystem functional groups, editado por David Keith; José R. Ferrer-Paris; Emily Nicholson; Richard T. Kingsford, 1-170. Gland, Suiça: IUCN, International Union for Conservation of Nature, 2020.
    Publicado
Pré-impressão
  1. Eliza Fragkopoulou; Térence Legrand; Juan David González-Trujillo; Jorge Assis. "The global biogeography of coastal oceanographic connectivity". 2024. https://doi.org/10.21203/rs.3.rs-4319669/v2.
    10.21203/rs.3.rs-4319669/v2
  2. Térence Legrand; Eliza Fragkopoulou; Juan David González-Trujillo; Jorge Assis. "The global biogeography of coastal oceanographic connectivity". 2024. https://doi.org/10.21203/rs.3.rs-4319669/v1.
    10.21203/rs.3.rs-4319669/v1
  3. Térence Legrand; Eliza Fragkopoulou; Lauren Vapillon; Lidiane Gouvêa; Ester A. Serrão; Jorge Assis. "Unravelling the role of oceanographic connectivity in intra-specific diversity of marine forests at global scale". 2023. https://doi.org/10.1101/2023.07.12.548579.
    10.1101/2023.07.12.548579
Tese / Dissertação
  1. "Potential consequences of future climate change for ecosystem structuring species". 2024. https://figshare.com/articles/thesis/Potential_consequences_of_future_climate_change_for_ecosystem_structuring_species/25306225.
    10.6084/m9.figshare.25306225
  2. Erzini, Karim; Costa, Barbara Horta e; Fragkopoulou, Eliza. "A global assessment of the effectiveness of different protection levels in marine protected areas of tropical waters, following a new categorization". Mestrado, 2015. http://hdl.handle.net/10400.1/7706.

Outros

Conjunto de dados
  1. Assis, Jorge; Fragkopoulou, Eliza. Global Marine Species Traits Database (marineSpeciesTraits). 2024.
    https://doi.org/10.6084/m9.figshare.27620676
Atividades

Apresentação oral de trabalho

Título da apresentação Nome do evento
Anfitrião (Local do evento)
2024 Global marine heatwaves pose risk to biodiversity across depths EuroMarine Open Science Day 2024
EuroMarine Network (Bologna, Itália)
2024 Potential Consequences of Climate Change on Marine Ecosystem Structuring Species Topics in Marine Biology Undergraduate Course
University of Algarve (Portugal)
2023 Global changes in species richness and community composition of ecosystem-structuring marine species under projected climate change Global changes in species richness and community composition of ecosystem-structuring marine species under projected climate change
ECCWO-5 International Symposium's. Effects of Climate Change on the World's Ocean (Bergen, Noruega)
2023 Global redistribution of cold-water coral species leading to widespread community turnover under climate change ISDSC 8- 8th International Symposium of Deep-Sea Corals
(Edinburgh, Reino Unido)
2023 Climate shaping genetic biodiversity of polar marine forests European Phycological Congress
(França)
2023 The MPA guide: A global meta-analysis of ecological outcomes of MPA protection levels and stages of establishment using the MPA guide The MPA guide: A global meta-analysis of ecological outcomes of MPA protection levels and stages of establishment using the MPA guide
(Vancouver, Canadá)
2023 Climate shaping genetic biodiversity of polar marine forests Climate shaping genetic biodiversity of polar marine forests
(Rhode Island, Estados Unidos)
2023 Potential consequences of future climate change on marine forests CCMAR seminar
CCMAR - Centro de Ciências do Mar (Faro, Portugal)
2022 Marine biodiversity exposed to prolonged and intense subsurface heatwaves ISMS - 8th International Symposium on Marine Sciences
Grand Canaria, Spain
2022 Major restructuring of marine forests' biodiversity under projected climate change ISMS - 8th International Symposium on Marine Sciences
(Grand Canaria, Espanha)
2021/10 Depth provides refugia for marine biodiversity during heatwave events. Early Career Biogeographers Conference
International Biogeography Society (Amsterdam, Países Baixos)
2021/07/22 Global species richness patterns of marine forests of large brown macroalgae. Annual meeting of the Phycological Society of America,
(Virtual)
2019/09 Dispersal mediated by ocean currents explains the genetic structure of marine forests 5th International Marine Connectivity Conference
(Aveiro, Portugal)
2016/07/15 Assessing the effectiveness of marine partially protected areas IMMR | International Meeting on Marine Research
(Peniche, Portugal)

Orientação

Título / Tema
Papel desempenhado 		Curso (Tipo)
Instituição / Organização
2024 - Atual PhD chapter: Improving SDM implementation and predicting the 4D distribution of marine commercial species
Coorientador
Centro de Ciências do Mar, Portugal
2023 - 2023 MSc thesis: Redistribution of sponge biodiversity under global climate change scenarios
Coorientador
Universidade do Algarve, Portugal
2022 - 2022 MSc thesis: Redistribution of cold-water coral biodiversity under global climate change scenarios
Coorientador
Universidade do Algarve, Portugal
2022 - 2022 MSc thesis: The global network of Marine Protected Areas is not ready to face projected climate change
Coorientador
Universidade do Algarve, Portugal
2021 - 2022 Internship on cold-water coral genetic data collection
Coorientador
Centro de Ciências do Mar, Portugal
2021 - 2022 Internship on assessing marine invasions: Posidonia versus Caulerpa
Coorientador
Centro de Ciências do Mar, Portugal
2021 - 2022 Biogeography of African algae patterns
Coorientador
Centro de Ciências do Mar, Portugal
2021 - 2021 Internship on marine forest genetic data collection
Orientador
Centro de Ciências do Mar, Portugal
2021 - 2021 Internship on marine heatwave mortality with depth
Coorientador
2021 - 2021 Internship on Sargassum golden tides data collection
Orientador
Centro de Ciências do Mar, Portugal
2020 - 2020 Internship on mangrove genetic data collection
Orientador
Centro de Ciências do Mar, Portugal
2020 - 2020 Internship on marine forest genetic data collection
Orientador
Universidade do Algarve, Portugal
2020 - 2020 Internship on marine forest genetic data collection
Orientador
Universidade do Algarve, Portugal

Entrevista (jornal / revista)

Descrição da atividade Jornal / Forum
2023 As oceans warm, marine heat waves push deep beneath the surface Mongabay
2023 Interview on the effects of subsurface marine heatwaves on biodiversity Publico

Entrevista / Programa (rádio / tv)

Programa Tema
2023/09/28 - 2023/09/28 Podcast on Clubhouse Climate Change and Extreme Events in the Ocean
2023 - 2023 National radio broadcasting Antenna 1 Subsurface marine heatwaves and their effects on biodiversity
2023 - 2023 National TV News program RTP Notícias Subsurface heatwaves and their effects on marine biodiversity

Membro de associação

Nome da associação Tipo de participação
2024 - Atual European Marine Board of Early Career Ocean Professionals (ECOPs)
2022 - Atual European Marine Science Network
2022 - Atual Group on Earth Observations Biodiversity (GEO BON)
2022 - Atual Ocean Decade initiative (U.N. Decade of Ocean Science for Sustainable Development)

Membro de comissão

Descrição da atividade
Tipo de participação 		Instituição / Organização
2021/07 - Atual Marine Biodiversity Observation Network - Group on earth observations
Membro
 Marine Biodiversity Observation Network - MBON, Portugal
2018 - Atual Marine Forests. Platform for free access to biodiversity distribution data of marine forests worldwide. World Wide Web electronic publication, CCMAR, http://www.marineforests.com
Membro
 Centro de Ciências do Mar, Portugal
Distinções

Outra distinção

2020 Doctorate Scholarship from the Portuguese Foundation of Science and Technology
Fundação para a Ciência e a Tecnologia, Portugal
2018 Bolsa de Investigação Mestre
Universidade do Algarve, Portugal
2011 Erasmus Placement Grant
Panepistemio Aigaiou, Grécia
2011 Principal´s Excellence Fund
University of Aberdeen, Reino Unido
2011 SAMS Research Bursary
Scottish Association for Marine Science, Reino Unido
2006 Distinction Grant of University of the Aegean, Greece
Panepistemio Aigaiou, Grécia